When Spaces is activated (i.e. you're viewing the overview window showing small versions of each Space), holding Command while using the arrow keys to navigate between the Spaces moves any window in the current Space into the new one.
For example, assume the top left Space is showing a Safari window, and that you have this Space selected in the Spaces overview. If you then press Command-Right Arrow, the Safari window will be moved to the top right Space. If there's a window open in that space (Mail maybe), and you press Command-Down, both the Mail window and the Safari window would end up in the bottom right window.
The only reason I can imagine they implemented this behavior would be if someone was trying to collect all open windows into one Space, maybe?
[robg adds: Based on a quick test, it looks like this new Command-Arrow behavior replaces this previous hint, which noted that you could press 'C' in the Spaces overview to collect all open windows. This no longer works in 10.6, nor does 'C' work with any combination of the modifier keys. It looks like there's no longer a 'collect all' shortcut.]
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